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Antonyms that are in the dictionary are marked in green. Antonyms that are not in the dictionary are marked in red.
Counts are ongoing, but NatureScot said initial observations from some of the main sites in Shetland suggested "significant breeding population declines" for great skuas.
The BAS suspects that migratory brown skuas from South America may have inadvertently brought the virus over to Bird Island, highlighting the risk that migration poses in spreading diseases across vast distances.